Beginning its journey in Rickmansworth, Hertfordshire in May 2021, it has excavated one of two tunnels at a depth of up to 80m (262ft).
A second, identical machine is expected to achieve its breakthrough in the coming weeks.
Rail minister Huw Merriman said it was a “groundbreaking moment ” and praised the dedication of the “450-strong team” that worked on it.
HS2 Ltd said about 17 people staffed an “underground factory” in shifts to keep it running constantly.
The tunnel was lined with 56,000 pre-cast concrete segments as the machine moved at an average speed of 16m (52ft) a day.
Florence, which has a diameter of 10.25m (34ft), will now be dismantled.
Some large parts will be reused in other tunnel boring machines (TBM) and other parts scrapped.
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