HES said it was working with Scottish Borders Council (SBC) to return the remains to their graves.
A spokesperson said: “This type of incident can occasionally happen in the setting of historic graveyards and, following best practice, we are working closely with SBC to ensure the discreet and respectful re-interment of the remains.”
An SBC spokesperson added: “We are liaising with HES colleagues regarding the monitoring of animal activity within the graveyard and how the issue can be addressed.”
The incident comes four years after two complete skeletons were uncovered at Jedburgh Abbey during renovations to the ramparts walkway.
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