The Animal Rising group said that two of its supporters had attacked the painting in a way that they claimed was “light-hearted”.
They pasted over the royal portrait with a picture of Wallace, a character from the Wallace and Gromit animations.
They were criticising the welfare standards of RSPCA “assured farm” status, with King Charles a patron of the RSPCA.
“With King Charles being such a big fan of Wallace and Gromit, we couldn’t think of a better way to draw his attention,” said Daniel Juniper, one of those involved.
The group said the posters had been attached to the glass by water.
The gallery said the incident had been over quickly and no harm had been done to the artwork.
The painting, with its strong red colours, drew much public attention when it was unveiled by King Charles in Buckingham Palace last month.
The RSPCA has been approached for a comment.
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